o Privatdozent in Biogeography
University of Basel, Switzerland 2014
Habilitation Title ‘Oldest Rainforest in Africa? Does the pattern of biological diversity correlate with environmental history’.
o Ph.D. in Molecular Genetics
University of Glasgow and Natural History Museum, London 2001-2006
Ph.D. Dissertation Title ‘Systematics and biogeography of amphibians of the Eastern Arc Mts’.
o M.Sc. Advanced methods in Biodiversity and Taxonomy
Imperial College and the Natural History Museum, London 1998-1999
Lecture course and MSc thesis on systematics of caecilian amphibians
o B.Sc. Zoology
University of Leeds 1995-1998
Lecture course and B.Sc. thesis on the behavioural ecology of sticklebacks